1. South China Botanical Garden; Chinese Academy of Sciences; Guangzhou 510650 China
2. Department of Land, Air, and Water Resources; University of California-Davis; One shields Avenue Davis CA 95616 USA
3. Xishuangbanna Tropical Botanical Garden; Chinese Academy of Sciences; Mengla 666303 China
4. Department of Environmental Sciences; East China Normal University; Shanghai 200062 China
5. Institute of Applied Ecology; Chinese Academy of Sciences; Shenyang 110164 China