1. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany Chinese Academy of Sciences Beijing 100093 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Jiangxi Province Key Laboratory of Watershed Ecosystem Change and Biodiversity, Institute of Life Science and School of Life Sciences Nanchang University Nanchang 330031 China
4. Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d'Histoire naturelle, CNRS, Sorbonne Université, EPHE Université des Antilles Paris 75005 France
5. Missouri Botanical Garden St. Louis Missouri 63166‐0299 USA