1. Plant Phylogenetics and Conservation Group, Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Kunming 650223 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Tibet Agriculture & Animal Husbandry University Nyingchi 860000 China
4. Australian Centre for Evolutionary Biology and Biodiversity and Sprigg Geobiology Centre, School of Biological Sciences University of Adelaide Adelaide SA 5005 Australia