1. Aquaculture Research Department, Production Engineering Division, Goto Field Station Japan Fisheries Research and Education Agency Goto‐shi Japan
2. Aquaculture Research Department, Aquatic Breeding Division, Kamiura Field Station Japan Fisheries Research and Education Agency Saeki‐shi Japan
3. Aquaculture Research Department, Aquatic Breeding Division, Tamaki Field Station Japan Fisheries Research and Education Agency Watarai‐gun Japan