1. Department of Biology, Faculty of Sciences Ferdowsi University of Mashhad Mashhad Iran
2. Zoological Innovations Research Department, Institute of Applied Zoology, Faculty of Sciences Ferdowsi University of Mashhad Mashhad Iran
3. Severtsov Institute of Ecology and Evolution RAS Moscow Russia
4. University of Gothenburg Department of Biology and Environmental Sciences, Systematics and Biodiversity Göteborg Sweden
5. Gothenburg Global Biodiversity Centre Gothenburg Sweden