1. Department of Ecology and Evolutionary Biology; Cornell University; Ithaca NY 14853 USA
2. Department of Food and Environmental Sciences/Microbiology; University of Helsinki; Viikki Biocenter; FIN-00014 Helsinki Finland
3. Swiss Federal Institute of Aquatic Science and Technology; Eawag; 8600 Dübendorf Switzerland
4. Department of Biological Statistics and Computational Biology; Cornell University; Ithaca NY 14853 USA