1. Hammersmith Medicines Research; Cumberland Avenue London NW10 7EW UK
2. Department of Cellular and Molecular Physiology; Institute of Translational Medicine, University of Liverpool; Crown Street Liverpool L69 3GE UK
3. Department of Gastroenterology and Hepatology, St. Olav's Hospital, and the Department of Cancer Research and Molecular Medicine; Norwegian University of Science and Technology; Trondheim Norway
4. Department of Pathology; Royal Liverpool and Broadgreen University Hospitals NHS Trust; Liverpool L7 8XP UK