1. Grice Marine Laboratory College of Charleston 205 Fort Johnson Rd Charleston SC 29412 USA
2. Department of Biology College of Charleston 66 George St. Charleston SC 29424 USA
3. Odum School of Ecology University of Georgia 130 E. Green St. Athens GA 30602 USA
4. NOAA/National Ocean Service Center for Coastal Environmental Health and Biomolecular Research 219 Fort Johnson Rd Charleston SC 29312 USA
5. GEOMAR Helmholtz‐Zentrum für Ozeanforschung Kiel Düsternbrooker Weg 20 D‐23105 Kiel Germany
6. Department of Fisheries Kagoshima University Shimoarata 3‐50‐20 Kagoshima City 890‐0056 Japan