1. School of Biological, Earth and Environmental Sciences Evolution & Ecology Research Centre UNSW Sydney Sydney New South Wales Australia
2. School of Biological Sciences Australian Centre for Ancient DNA (ACAD) University of Adelaide Adelaide South Australia Australia
3. Biosciences Institute Newcastle University Newcastle upon Tyne UK
4. Behavioural Ecology and Ecophysiology Group Department of Biology University of Antwerp Wilrijk Belgium
5. Section of Amphibians and Reptiles Carnegie Museum of Natural History Pittsburgh Pennsylvania USA