1. School of Biosciences University of Sheffield Western Bank Sheffield S10 2TN UK
2. Institute of Evolutionary Biology, School of Biological Sciences The University of Edinburgh, Ashworth Laboratories Charlotte Auerbach Road Edinburgh EH9 3FL UK
3. Wellcome Sanger Institute Wellcome Genome Campus, Hinxton Cambridge CB10 1SA UK
4. Department of Marine Sciences University of Gothenburg S‐405 30 Gothenburg Sweden