1. College of Life and Environmental Sciences Hangzhou Normal University Hangzhou 311121 China
2. CAS Plant Germplasm and Genomics Center Germplasm Bank of Wild Species Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
3. CAS Key Laboratory for Plant Biodiversity and Biogeography of East Asia Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
4. Department of Life Sciences Natural History Museum London SW7 5BD UK
5. Key Laboratory of Tropical Plant Resources and Sustainable Use Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Menglun Yunnan 666303 China
6. Center of Integrative Conservation Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Menglun Yunnan 666303 China