1. Centre for African Ecology School of Animal, Plant and Environmental Sciences University of the Witwatersrand Wits 2050 South Africa
2. Institute of Biodiversity, Animal Health and Comparative Medicine University of Glasgow Glasgow G12 8QQ UK
3. CEFE CNRS Univ. Montpellier Univ. Paul Valéry Montpellier 3 EPHE IRD Montpellier France
4. School of Animal, Plant and Environmental Sciences University of the Witwatersrand Wits 2050 South Africa
5. Okavango Research Institute University of Botswana Maun Botswana
6. Resource Ecology Group Wageningen University Wageningen 6708 PB The Netherlands