1. Department of Biology, Faculty of Science; Shinshu University; Matsumoto Japan
2. Institute of Mountain Science; Shinshu University; Matsumoto Japan
3. Faculty of Geo-environmental Science; Rissho University; Kumagaya Japan
4. Division of Mountain and Environmental Science, Interdisciplinary Graduate School of Science and Technology; Shinshu University; Matsumoto Japan
5. Akkeshi Marine Station, Field Science Center for Northern Biosphere; Hokkaido University; Akkeshi Hokkaido Japan
6. Division of Developmental Science, Graduate School for International Development and Cooperation; Hiroshima University; Higashi-Hiroshima Japan
7. Department of Entomology; University of Wisconsin-Madison; Madison Wisconsin USA