1. Infection and Immunity Program and Department of Biochemistry and Molecular Biology Biomedicine Discovery Institute Monash University Clayton VIC 3800 Australia
2. Australian Research Council Centre of Excellence for Advanced Molecular Imaging Monash University Clayton VIC 3800 Australia
3. Department of Microbiology and Immunology Peter Doherty Institute for Infection and Immunity University of Melbourne Parkville VIC 3010 Australia
4. Division of Infection and Immunity Cardiff University, School of Medicine Heath Park Cardiff CF14 4XN UK