1. Innovative Center of Molecular Genetics and Evolution School of Life Sciences, Guangzhou University Guangzhou 510006 China
2. The Innovative Academy of Seed Design, Key Laboratory of Soybean Molecular Design Breeding, Northeast Institute of Geography and Agroecology the Chinese Academy of Sciences Harbin 150000 China
3. Key Laboratory of South China Agricultural Plant Molecular Analysis and Genetic Improvement South China Botanical Garden, the Chinese Academy of Sciences Guangzhou 510650 China
4. National Key Laboratory of Crop Genetics and Germplasm Enhancement Jiangsu Collaborative Innovation Center for Modern Crop Production, Nanjing Agricultural University Nanjing 210095 China