1. Research Centre on Animal Cognition Center for Integrative Biology CNRS (UMR 5169) University of Toulouse Toulouse France
2. Laboratoire Evolution et Diversité Biologique CNRS IRD (UMR 5174) University of Toulouse Toulouse France
3. Evolution, Genomes, Behavior and Ecology CNRS IRD (UMR 9191 University Paris Saclay Gif‐sur‐Yvette France
4. College of Animal Sciences (College of Bee Science) Fujian Agriculture and Forestry University Fuzhou China
5. Institut Universitaire de France (IUF) Paris France