1. INRA; LBLGC; EA 1207; University of Orléans; Orléans F-45067 France
2. UMR547 PIAF; INRA; Clermont-Ferrand F-63100 France
3. UMR547 PIAF; Clermont Université, Université Blaise-Pascal; Clermont-Ferrand F-63000 France
4. Department of Biology; Centre of Excellence, Plant and Vegetation Ecology (PLECO); University of Antwerp; Wilrijk B-2610 Belgium