1. Department of Zoology & Entomology University of Pretoria Hatfield South Africa
2. Department of Terrestrial Invertebrates National Museum Bloemfontein 36 Aliwal Street Bloemfontein 9300 South Africa
3. Laboratory of Zoology and Aquatic Ecology – LAZOEA University of los Andes Bogotá Colombia
4. Programa de Pós‐Graduação em Ecologia, Conservação e Manejo da Silvestre Universidade Federal de Minas Gerais Belo Horizonte MG Brazil