1. Muséum National d'Histoire Naturelle; Institut de Systématique Evolution Biodiversité; Herbier National; Case Postale 39, 12 rue Buffon 75004 Paris France
2. Department of Biology, School of Life Sciences; East China Normal University; 3663 Zhong Shan North Road Shanghai 200062 China
3. Herbario QCA, Escuela de Ciencias Biológicas; Pontificia Universidad Católica del Ecuador; Apartado 17-01-2184 Quito Ecuador