1. School of Biology University of St Andrews, College Gate St AndrewsKY16 9AJUK
2. Department of Collective Behavior Max Planck Institute for Animal Behavior Radolfzell Germany
3. Centre for the Advanced Study of Collective Behaviour University of Konstanz Universitätsstraße 1078464Konstanz Germany
4. Department of Biology University of Konstanz Universitätsstraße 1078464Konstanz Germany
5. Department of Biology Pacific University 2043 College Way Forest Grove OR97116USA