1. Center for Computational Biology College of Biological Sciences and Technology Beijing Forestry University Beijing 100083 China
2. Beijing Key Laboratory of Ornamental Plants Germplasm Innovation and Molecular Breeding National Engineering Research Center for Floriculture College of Landscape Architecture Beijing Forestry University Beijing 100083 China
3. Department of Biostatistics Rutgers University New Brunswick NJ 08901 USA
4. Arvalis Institut du Végétal 3‐5 Rue Joseph et Marie Hackin 75116 Paris France
5. Center for Statistical Genetics Departments of Public Health Sciences and Statistics Pennsylvania State University Hershey PA 17033 USA