1. Department of Life Science National Taiwan Normal University Taipei 106 Taiwan
2. Biodiversity Program, Taiwan International Graduate Program Academia Sinica Taipei 115 Taiwan
3. Biodiversity Research Center Academia Sinica Taipei 115 Taiwan
4. Natural History Museum of Denmark University of Copenhagen Copenhagen 2100 Denmark
5. Institute of Ecology and Evolutionary Biology National Taiwan University Taipei 106 Taiwan
6. White Sea Biological Station Biological Faculty of Moscow State University Moscow 119899 Russia
7. Marine Biological Section, Department of Biology University of Copenhagen Copenhagen 2100 Denmark