1. Department of Fisheries, Faculty of Agriculture and Natural Resources, Science and Research Branch; Islamic Azad University; Tehran Iran
2. Department of Aquatic Animal Health, Faculty of Veterinary Medicine; University of Tehran; Tehran Iran
3. Freshwater Fish Group and Fish Health Unit, School of Veterinary and Life Sciences; Murdoch University; Murdoch Western Australia Australia
4. International Sturgeon Research Institute; Rasht Iran