1. Department of Microbiology and Immunology; Hirosaki University Graduate School of Medicine; Hirosaki Aomori Japan
2. Laboratory of Food Safety Science; Department of Veterinary Medicine; Faculty of Agriculture; Iwate University; Morioka Iwate Japan
3. Department of Basic Veterinary Science; United Graduate School of Veterinary Medicine; Gifu University; Gifu Japan
4. Laboratory of Zoonoses; Kitasato University School of Veterinary Medicine; Towada Aomori Japan
5. Laboratory of Veterinary Anatomy and Cell Biology; Department of Veterinary Medicine; Faculty of Agriculture; Iwate University; Morioka Iwate Japan