1. Department of Zoology; University of Otago; Dunedin New Zealand
2. Institute of Veterinary, Animal and Biomedical Sciences; Massey University; Palmerston North New Zealand
3. Department of Zoology; University of Oxford; Oxford UK
4. Charles Perkins Centre; Faculty of Veterinary Sciences, and School of Biological Sciences; University of Sydney; Sydney Australia
5. Institute of Natural Sciences; Massey University; Auckland New Zealand