1. Department of Integrated Biosciences, Graduate School of Frontier Sciences, The University of Tokyo Chiba Japan
2. Earth and Sustainability Science Research Centre, School of Biological, Earth and Environmental Sciences, The University of New South Wales Sydney NSW Australia
3. University and Jepson Herbaria, and Department of Integrative Biology, University of California Berkeley Berkeley CA USA
4. Department of Biological Sciences, Graduate School of Science, The University of Tokyo Tokyo Japan
5. Atmosphere and Ocean Research Institute, The University of Tokyo Chiba Japan