1. Centre for Environmental Biology and Ecosystem Studies; National Institute for Environmental Studies; 16-2 Onogawa Tsukuba Ibaraki 305-8506 Japan
2. Department of Forest Genetics; Forestry and Forest Products Research Institute; 1 Matsunosato Tsukuba Ibaraki 305-8687 Japan
3. Institute of Boninology; Miyanohamamichi Chichijima Ogasawara Tokyo 100-2101 Japan
4. Graduate School of Agriculture; Kyoto University; Kitashirakawa Oiwake-cho Sakyo-ku Kyoto 606-8502 Japan
5. Primate Research Institute; Kyoto University; 41-2 Kanrin Inuyama Aichi 484-8506 Japan
6. Wildlife Research Centre; Kyoto University; 2-24 Tanaka-Sekiden-cho Sakyo-ku Kyoto 606-8203 Japan