1. Department of Ecology and Evolutionary Biology; University of California Los Angeles; 621 Charles E. Young Drive South Los Angeles California 90095 USA
2. Key Laboratory of Tropical Forest Ecology; Xishuangbanna Tropical Botanical Garden; Chinese Academy of Sciences; Mengla Yunnan 66303 China
3. State Key Laboratory for Conservation and Utilization of Subtropical Agro-bioresources, and College of Forestry; Guangxi University; Nanning Guangxi 530005 China