1. Center for Ecological Research; Kyoto University; Hirano 2-509-3 Otsu 5202113 Japan
2. Entomological Laboratory; Graduate School of Life and Environmental Science; Osaka Prefecture University; Gakuencho1-1 Nakaku Sakai 5998531 Japan
3. Department of Animal Sciences; Teikyo University of Science & Technology; 2525 Yatsusawa Uenohara Yamanashi 4090193 Japan