1. Ocean and Earth Science, National Oceanography Centre, Univ. of Southampton; Southampton SO14 3ZH UK
2. Dept of Integrative Biology; Oregon State Univ.; Corvallis OR 97331 USA
3. Dept of Ecology and Evolutionary Biology; Long Marine Laboratory, Univ. of California; Santa Cruz CA 95060 USA
4. Centro de Estudios Avanzados en Zonas Áridas (CEAZA) and Univ. Católica del Norte; CP. 1781421 Coquimbo Chile