1. USDA, APHIS, WS, National Wildlife Research Center, 4101 LaPorte Avenue; Fort Collins CO 80521 U.S.A.
2. Pacific Cooperative Studies Unit; University of Hawai'i, 3190 Maile Way; Honolulu HI 96822 U.S.A.
3. U.S. Geological Survey, Pacific Island Ecosystems Research Center, PO Box 369; Makawao HI 96768 U.S.A.