1. Department of Evolution, Ecology and Behaviour, Institute of Integrative Biology; University of Liverpool; Liverpool L69 7ZB U.K.
2. Evolutionary Biology; Bielefeld University; 33615 Bielefeld Germany
3. Evolution and Ecology Research Centre, School of Biological, Earth and Environmental Sciences; University of New South Wales; Sydney 2052 Australia
4. Division of Ecology and Evolution, Research School of Biology; The Australian National University; 2601 Canberra Australia
5. Institute of Zoology; University of Graz; Graz 8010 Austria