1. Population Biology and Conservation Biology/Department of Ecology and Genetics; Evolutionary Biology Centre; Uppsala University; Norbyvägen18D 752 36 Uppsala Sweden
2. Department of Animal Biology, Plant Biology and Ecology; Faculty of Science; University of A Coruña; Campus da Zapateira 15071 A Coruña Spain
3. Department of Clinical Sciences; Computational Genetics Section; Swedish University of Agricultural Sciences; 756 51 Uppsala Sweden
4. Department of Animal Biology; University of Barcelona; Diagonal 643 08028 Barcelona Spain