1. INRA, Agrocampus-Ouest, Université de Rennes 1; UMR1099 IGEPP Rennes Cedex France
2. School of Biological, Biomedical and Environmental Sciences; University of Hull; Hull U.K.
3. INRA-UMR Centre de Biologie et de Gestion des Populations, CBGP, (INRA/IRD/CIRAD/Montpellier SupAgro); Montferrier-sur-Lez France