1. LARG; Department of Zoology; University of Cambridge; Cambridge UK
2. CEFE-CNRS; Montpellier Cedex 5 France
3. Department of Zoology; The Edward Grey Institute; University of Oxford, Oxford UK
4. Institute of Evolutionary Biology; School of Biological Sciences; University of Edinburgh; Edinburgh UK
5. Institute of Zoology; Zoological Society of London; London UK