1. Biology Department; 1878 Campus Delivery; Colorado State University; Fort Collins CO 80523 U.S.A.
2. Department of Biology; Program in Ecology, Evolution, and Conservation Biology; 1664 N. Virginia Street, University of Nevada Reno NV 89557 U.S.A.
3. Department of Environmental Science and Policy; One Shields Avenue; University of California; Davis CA 95616 U.S.A.
4. Department of Evolution and Ecology; Center for Population Biology; One Shields Avenue; University of California; Davis CA 95616 U.S.A.