1. Laboratory of Protozoology Institute of Evolution and Marine Biodiversity Ocean University of China Qingdao 266003 China
2. Key Laboratory of Aquatic Biodiversity and Conservation of Chinese Academy of Sciences Institute of Hydrobiology Chinese Academy of Sciences Wuhan 430072 China
3. College of Life Sciences Capital Normal University Beijing 100048 China
4. Department of Biological Sciences Boise State University Boise Idaho 83725 USA