1. Australian Research Council Centre of Excellence for Environmental Decisions; School of Biological Sciences; The University of Queensland; Brisbane Qld 4072 Australia
2. Australian Research Council Centre of Excellence for Coral Reef Studies; School of Biological Sciences; The University of Queensland; Brisbane Qld 4072 Australia
3. Marine Ecology Research Centre; Southern Cross University; Lismore NSW 2480 Australia
4. National Marine Science Centre; Southern Cross University; Coffs Harbour NSW 2450 Australia
5. Marine Ecology Research Centre; Southern Cross University; Coffs Harbour NSW 2450 Australia