1. INRA, UMR Eco&Sols, INRA-IRD-Cirad-SupAgro; 2 Place Viala 34060 Montpellier France
2. Institute of Biotechnology, Vietnam Academy of Science and Technology; 18 Hoang Quoc Viet Road Hanoi Vietnam
3. Cirad, UMR17, Intertryp, Cirad-IRD; TA-A17/G, Campus International de Baillarguet 34398 Montpellier Cedex 5 France
4. UM2, UMR 5236, CPBS, CNRS-UM1-UM2; 1919 route de Mende 34293 Montpellier Cedex 5 France