1. Key Laboratory for Bio-Resources and Eco-Environment; College of Life Sciences, Sichuan University; Chengdu 610065 China
2. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650201 China
3. College of Life Sciences; South China Agricultural University; Guangzhou 510642 China
4. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden; Chinese Academy of Sciences; Guangzhou 510650 China