1. Division of Agricultural and Environmental Sciences, Gateway Building, Sutton Bonington CampusUniversity of Nottingham Leicestershire LE12 5RD UK
2. Sustainable Soils and Grassland Systems DepartmentRothamsted Research, West Common, Harpenden Hertfordshire AL5 2JQ UK
3. Metabolic Biology, John Innes CentreNorwich Research Park Norwich NR4 7UH UK