1. College of Environmental Science and Engineering Ocean University of China Qingdao 266100 China
2. Laboratory for Marine Ecology and Environmental Science Pilot National Laboratory for Marine Science and Technology (Qingdao) Qingdao 266237 China
3. Key Laboratory of Marine Environment and Ecology Ministry of Education Qingdao 266100 China
4. Fourth Institute of Oceanography Ministry of Natural Resources Beihai 536002 China