1. School of Karst Science Guizhou Normal University Guiyang China
2. Department of Anthropology University of Illinois Urbana Illinois USA
3. Ministry of Education Key Laboratory for Ecology of Tropical Islands, Key Laboratory of Tropical Animal and Plant Ecology of Hainan Province, College of Life Sciences Hainan Normal University Haikou China
4. Bawangling Branch Hainan Tropic Rainforest National Park Administration China