1. State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences; Kunming Yunnan China
2. Graduate School of Chinese Academy of Sciences; Beijing China
3. College of Animal Science and Technology; Sichuan Agriculture University; Ya'an Sichuan China
4. Department of Anthropology; Washington University in St. Louis; St. Louis Missouri USA
5. Institute of Eastern-Himalaya Biodiversity Research; Dali University; Dali Yunnan China