1. School of Life Sciences Institute of Evolution and Ecology Central China Normal University Wuhan China
2. State Key Laboratory of Integrated Management of Pest Insects and Rodents in Agriculture Chinese Academy of Sciences Institute of Zoology Beijing China
3. State Key Laboratory of Grassland Agro‐ecosystem Institute of Innovation Ecology & College of Life Sciences Lanzhou University Lanzhou China
4. College of Life Sciences Qufu Normal University Qufu China