1. Ministry of Education Key Laboratory for Ecology of Tropical Islands College of Life Sciences Hainan Normal University Haikou China
2. State Forestry Administration of China Key Laboratory for Biodiversity Conservation in Mountainous Areas of Southwest Karst School of Life Sciences Guizhou Normal University Guiyang China
3. Ecologie Systématique Evolution Université Paris‐Sud CNRS AgroParisTech Université Paris‐Saclay Orsay Cedex France