1. Marine College, Shandong University Weihai China
2. Institute of Ecology and Biodiversity, Shandong University Jinan China
3. The Kyoto University MuseumKyoto University Kyoto Japan
4. College of Life Science, Guangzhou University Guangzhou China
5. Institute of Tropical Biosciences and Biotechnology, Chinese Academy of Tropical Agricultural Sciences Haikou China