1. State Key Laboratory of Grassland Agro‐ecosystems, College of Ecology & School of Life Sciences Lanzhou University Lanzhou China
2. Institute of Evolution and Ecology, College of Life Sciences Central China Normal University Wuhan China
3. College of Ecology and Environmental Science Yunnan University Kunming China
4. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology and Institute of Biodiversity Yunnan University Kunming China