1. Key Laboratory of Zoological Systematics and Evolution Institute of Zoology Chinese Academy of Sciences Beijing China
2. University of Chinese Academy of Sciences Beijing China
3. Key Laboratory of Forest Ecology in Tibet Plateau of Ministry of Education Institute of Tibet Plateau Ecology Tibet Agriculture & Animal Husbandry University Nyingchi China
4. Key Laboratory of Animal Physiology, Biochemistry and Molecular Biology of Hebei Province, College of Life Sciences Hebei Normal University Shijiazhuang China
5. Center for Excellence in Animal Evolution and Genetics Chinese Academy of Sciences Kunming China