1. Department of Natural Resources Science University of Rhode Island Kingston RI USA
2. Rubenstein School of Environment and Natural Resources University of Vermont Burlington VT USA
3. Department of Fish, Wildlife, and Conservation Biology and Graduate Degree Program in Ecology Colorado State University Fort Collins CO USA
4. U.S. Geological Survey Fort Collins Science Center Fort Collins CO USA
5. Colorado Parks and Wildlife Fort Collins CO USA
6. U.S. Geological Survey, Washington Cooperative Fish and Wildlife Research Unit, School of Environmental and Forest Sciences & School of Aquatic and Fishery Sciences University of Washington Seattle WA USA